That's partly because the organs that did the work of breathing -- either gills or lungs -- don't fossilize well.
这在一定程度上是因为负责呼吸的器官——鳃或肺——没有很好地变成化石。
英语百科
Work of breathing
↑ Force = Pressure x Area, and Distance = Volume / Area. When both refer to the same area, Force x Distance = (Pressure x Area) x (Volume/Area) = Pressure x Volume