An arthropod (from Greek arthro-, joint + podos, foot) is an invertebrate animal having an exoskeleton (external skeleton), a segmented body, and jointed appendages (paired appendages). Arthropods form the phylum Arthropoda, and include the insects, arachnids, myriapods, and crustaceans. Arthropods are characterized by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. The rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Their versatility has enabled them to become the most species-rich members of all ecological guilds in most environments. They have over a million described species, making up more than 80% of all described living animal species, some of which, unlike most animals, are very successful in dry environments.