This cooling trend continued into the early Miocene Epoch and expanded areas of semi-arid climate with more shrub lands, dunes, and savannas, gradually replacing the tall trees that these animals preferred.

这种降温趋势一直持续到中新世早期， 半干旱气候区域扩大，灌木丛、沙丘和稀树草原逐渐增多， 逐渐取代了这些动物喜欢的高大树木。