Central to their compassionate and complex society are the matriarchs.

大象慈悲且复杂的社会核心是其女族长们。

This beneficial effect was only seen when the matriarchs lived nearby.

只有在年长女性生活在附近时，这种有益作用才会显现出来。

Elephants live in complex, tightly-knit, multigenerational societies, led by older females known as matriarchs.

大象生活在复杂、紧密的多代社会中，由年长的母象领导。

And, finally, the royal family saying goodbye to its matriarch after a period of mourning.

最后，让我们来了解一下英国皇室在不久前的哀悼之后为前英国女王举行的告别仪式。

Researchers concluded that the matriarchs likely remembered where sources of water could be located.

研究人员得出结论，母象可能记得水源的位置。

And of course, a good matriarch would always keep her young away from such a threat.

当然，一个优秀的女族长一直都会保护小象，让它们免受如此的威胁。

An older matriarch, the leader of the herd, is accompanied by her daughters and other female relatives.

年长的母象王，是象群的首领。陪伴在旁的有女儿们和其他母象亲戚。

They had names, like David Greybeard, the first one to lose his fear; Old Flo, that amazing matriarch.

他们有名字，像是灰胡子大卫，他是第一隻放下恐惧的黑猩猩；老芙洛，那了不起的女家长。

Normally herds are led by an older matriarch who would know how to get Goa out of this maze.

通常象群由一头年长的雌象带领，她知道如何让果阿走出迷宫。

In 2003, Eleanor, an elephant matriarch, collapsed.

2003年，大象女族长埃莉诺倒下了。